Dell OptiPlex 990 USFF vs. HP ProDesk 400 G2.5 SFF
Comparison and Differences
Differences between Dell OptiPlex 990 USFF and HP ProDesk 400 G2.5 SFF
If you are looking for a more modern desktop, then you should go with the HP ProDesk 400 G2.5 SFF which was released in 2014. The Dell OptiPlex 990 USFF is older and entered production in 2011.
If you are looking for the smaller form factor the Dell OptiPlex 990 USFF is the right one for you. It is smaller than the HP ProDesk 400 G2.5 SFF and will fit more easily under your desk.
Processor
Usually the newer the generation of the CPU, the better the performance and efficiency is. If your budget allows it, you should go with the HP ProDesk 400 G2.5 SFF as it is running a newer generation CPU. This is not to say that Dell OptiPlex 990 USFF is bad - it can still run some powerful CPUs, albeit being an older generation.
Memory
Both models have have equal amount of ram slots (2 RAM). Usually two RAM slots are more than enough for most cases. However, the count should be taken into account when planning how much and what combination of RAM you are going to need for your system.
The HP ProDesk 400 G2.5 SFF is using the faster 1600 MT/s RAM. The Dell OptiPlex 990 USFF is slower by supporting RAM speeds of up to 1333 MT/s. Additionally, if you will be needing a lot of RAM, better go for the HP ProDesk 400 G2.5 SFF as it has the ability to run 16 GB of RAM. The Dell OptiPlex 990 USFF can do a bit less at max 8 GB. Most of the times, this should be sufficient for the majority of users.
Ports
In total the HP ProDesk 400 G2.5 SFF has 8 USB ports. The Dell OptiPlex 990 USFF is inferior in this regard and packs 7 USB ports. In any case, if you miss USB ports, you can always purchase a USB hub. None of the desktop models has a USB 3.1 port (10Gb/s), so if you need a fast connection to your peripherals, like external Solid State Drive, you will need to look elsewhere.
The DisplayPort is usually the more popular video port used with desktops and laptops, while the HDMI is used predominantly in consumer electronics. That's why, it is good to see that both of the models come with a DisplayPort.
None of the models has an HDMI port. If your monitor supports only the HDMI interface, you will need to buy an adapter. Such adapters are cheap so nothing to worry about.
SATA ports are placed on the system board and it is where you connect your HDD, SSD and Optical Disk Drives. More SATA slots will allow you to have more drives running at the same time. In this regard the HP ProDesk 400 G2.5 SFF is better equipped as it comes with 3 SATA slot(s), while the Dell OptiPlex 990 USFF has 2 slot(s).
M.2 SSD interface allows you to transfer data with higher speeds compared to the old SATA interface. Unfortunately, neither the Dell OptiPlex 990 USFF, nor the HP ProDesk 400 G2.5 SFF has one. However, you can still use a SATA SSD drive which should give you satisfactory performance.
Make sure that the available M.2 slot supports the PCIe (also called NVMe) interface. This is important as there are M.2 slots which support only the inferior SATA III interface. For comparison, the SATA III interface has max speeds of 6Gb/s, while the PCIe 3.0 x4 will support speeds up to 32Gb/s!
Power Supply
Having a Power Supply Unit with high power rating is important, if you are going to use components which require extra power. The HP ProDesk 400 G2.5 SFF comes with a decent 180 Watt PSU, but the Dell OptiPlex 990 USFF has a more powerful one rated at 200 Watts. Choosing either one depends on your specific requirements and use case. Also don't forget that some models might have more than one PSU option - try to always get the most powerful one, especially if you plan to upgrade to a powerful GPU.